Three months have passed since the previous series, and it is now spring 1939. Some big changes are happening at Skeldale, and everyone has to learn how to adjust. Tristan is now a qualified vet, while James enters a new stage of his life with Helen and at the practice. After being made joint business partner by Siegfried, James pushes to take on more responsibility via the Ministry of Agriculture’s new bovine TB testing scheme, but its unexpected challenges risk pushing him to breaking point. And as a potential world war looms, all the Skeldale family have to consider their purpose in Darrowby and beyond.

Everyone is trying to get along as best they can in the face of changes at Skeldale House. Helen has moved into the bedsit upstairs with James and though they’re happy, she’s unsure how to integrate herself into the rest of the house. No-one sees this more than Mrs Hall, who is keen to find a way to bring everyone together around the dinner table.

James struggles to get Siegfried to let him take the reins now that he’s junior partner at the practice. With Helen’s help, James learns how much money Skeldale could make from the Ministry of Agriculture’s lucrative bovine TB testing scheme, but Siegfried believes the MAG’s scheme is more trouble than it’s worth and risks damaging their relationships with farmers. He digs his heels in and a rift forms between the two vets. It takes Helen, Mrs Hall and a case of some mysteriously poisoned calves at farmer Kate Billings’ to help James and Siegfried see that they’re better when they work together.

A lackadaisical Tristan thinks that James being his boss will make his life even more of a breeze. He learns that he needs to step up in his role at the practice and take it more seriously, when the dog of a rival is rushed into the surgery in an emergency and no-one is there to help him.

This episode originally aired on September 22 and is on again at 5.40pm today (September 24) on Channel 5